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698 Full-time PhD programmes in Biology

2
Princeton University4,4(35)
Princeton, United States Top 0.5% in Worldwide

Ecology and Evolutionary Biology

The Ecology and Evolutionary Biology program from Princeton University is designed to develop both the breadth and depth of understanding that will enable graduates of the department to respond to future advances in the field. 

Ph.D. On-campus
5 yearsTuition not available
University of Cyprus4,1(95)
Nicosia, Cyprus Top 8% in Worldwide

Biodiversity and Ecology

The Ph.D. Degree in Biodiversity and Ecology from University of Cyprus aims at providing students with high quality education of international standing, so that that they can develop a deep understanding of their science and a versatile thinking, something critical for their professional career in the modern environment.

Ph.D. On-campus
4 years$4,495/yr
West Virginia University4,3(9)
Morgantown, United States Top 15% in Worldwide

Biology

The Department of Biology at West Virginia University offers graduate courses and research that lead to MS and PhD degrees in the areas of ecology, bioinformatics, genetics, genomics and evolutionary biology, biochemistry and molecular biology, neurobiology, behavior and physiology, plant sciences, cell and developmental biology, biology education and forensic biology.

Ph.D. On-campus
5 years$30,024/yr
University of British Columbia4,2(195)
Vancouver, Canada Top 1% in Worldwide

Biochemistry and Molecular Biology

The Biochemistry and Molecular Biology Graduate Program at University of British Columbia provides advanced, research-based education that will prepare students for a career in academic, industrial, or professional positions in British Columbia and beyond.  

Ph.D. On-campus
6 years$7,074/yr
German Cancer Research Center (DKFZ) in Heidelberg3,0(3)
Heidelberg, Germany

Cell Biology and Tumor Biology

The Research Program (FSP-A) at German Cancer Research Center (DKFZ) in Heidelberg concentrates the basic Cell Biology and Tumor Biology-focused research groups at the Center. 

Ph.D. On-campus
3 yearsFree
The University of Alabama at Birmingham3,8(11)
Birmingham, United States Top 9% in Worldwide

Biochemistry and Structural Biology

Biochemistry and Structural Biology (BSB) is one of eight interdisciplinary PhD themes within Graduate Biomedical Sciences at The University of Alabama at Birmingham.  Students select a theme upon application but have access to faculty and courses from across GBS, allowing for flexibility in both research and academics.

Ph.D. On-campus
7 years$12,975/yr
University of Pennsylvania4,3(22)
Philadelphia, United States Top 0.5% in Worldwide

Cell and Molecular Biology - Cancer Biology

The Cell and Molecular Biology - Cancer Biology Pprogram at the University of Pennsylvania is designed to train students in the core principles and experimental approaches within cancer biology and to empower their ability to conduct hypothesis-driven research to better understand the fundamental mechanisms that cause and define the hallmarks of cancer.

Ph.D. On-campus
5 years$44,792/yr
Ludwig-Maximilians-University (LMU) Munich4,1(100)
München, Germany Top 1% in Worldwide

Life Science - From Molecules to Systems

The Graduate School Life Science Munich (LSM) offers an international doctoral program in life sciences to outstandingly motivated and qualified graduates. We offer a Life Science - From Molecules to Systems degree at Ludwig-Maximilians-University (LMU) Munich. 

Ph.D. On-campus
3 yearsFree
Wake Forest University3,5(2)
Winston-Salem, United States Top 7% in Worldwide

Molecular and Cellular Biosciences

The Molecular and Cellular Biosciences (MCB) program at Wake Forest University allows students to enter graduate school with the freedom and flexibility to explore several programs before committing to a specific research topic. A range of mentors can be selected whose research spans basic and translational fields of cutting-edge molecular and cellular biology.

Ph.D. On-campus
5 years$38,650/yr
Wayne State University3,7(3)
Detroit, United States Top 8% in Worldwide

Cancer Biology

In the Cancer Biology program at Wayne State University you will find a strong interdisciplinary graduate curriculum with a major focus on the biology of cancer, and opportunities to interact with clinicians engaged in cancer diagnosis and treatment.

Ph.D. On-campus
7 years$25,174/yr
University of Virginia4,3(35)
Charlottesville, United States Top 3% in Worldwide

Experimental Pathology

The Molecular and Cellular Basis of Disease (MCBD) graduate program at University of Virginia confers the PhD in Experimental Pathology.  

Ph.D. On-campus
4 years$34,302/yr
Georgia Institute of Technology4,4(50)
Atlanta, United States Top 2% in Worldwide

Biology

The purpose of our PhD program in Biology at Georgia Institute of Technology is to prepare highly qualified scientists who have excellent up-to-date training in the fields of biology or bioinformatics.

Ph.D. On-campus
5 years$4,687/yr
Texas A&M University4,1(17)
College Station, United States Top 3% in Worldwide

Biology

The Ph.D. in Biology at Texas A&M University combines in-depth laboratory-conducted research with rigorous scholarly exploration of molecular, cellular, and systems principles across microbial, plant, and animal biology.

Ph.D. On-campus
5 years$16,217/yr
McMaster University4,5(68)
Hamilton, Canada Top 2% in Worldwide

Chemical Biology

In its broadest term, chemical biology is defined as the application of chemical principles to biochemistry and the medical sciences. This Chemical Biology degree is offered at McMaster University.

Ph.D. On-campus
4 years$3,009/yr
Uppsala University4,2(113)
Visby, Sweden Top 2% in Worldwide

Biology

Doctoral education in Biology at Uppsala University is offered at the Evolutionary Biology Centre (EBC, Department of Ecology and Genetics and Department of Organismal Biology) and Uppsala Biomedical Centre (BMC, Department of Cell and Molecular Biology).

Ph.D. On-campus
4 yearsFree
Kent State University4,2(18)
Kent, United States Top 15% in Worldwide

Biological Sciences - Ecology and Evolutionary Biology

The Biological Sciences - Ecology and Evolutionary Biology degree from Kent State University provides opportunities to study in areas such as animal behavior, entomology, limnology, microbial ecology, ornithology, systems ecology, systematic and evolutionary biology, environmental physiology, vertebrate ecology and population and community ecology.

Ph.D. On-campus
4 years$25,091/yr
German Cancer Research Center (DKFZ) in Heidelberg3,0(3)
Heidelberg, Germany

Functional and Structural Genomics

Cancer arises when genes in a cell are changed in such a way that they cause the cell to divide uncontrolled. For this to happen, however, a multitude of specific changes have to coincide. We offer a Functional and Structural Genomics degree at the German Cancer Research Center (DKFZ) in Heidelberg. 

Ph.D. On-campus
3 yearsFree
Tufts University4,1(72)
Medford, United States Top 4% in Worldwide

Genetics, Molecular and Cellular Biology

The Genetics, Molecular and Cellular Biology Program (GMCB) at Tufts University offers strong interdisciplinary training across a spectrum of fundamental biomedical areas, including classical and molecular genetics; structural & chemical Biology; and cell & developmental biology.

Ph.D. On-campus
5½ years$19,326/yr
The University of Iowa4,0(4)
Iowa City, United States Top 5% in Worldwide

Cell and Developmental Biology

The Cell and Developmental Biology (CDB) Program at The University of Iowa is housed in the Department of Anatomy and Cell Biology and includes faculty mentors from other departments. It is part of the larger Biomedical Sciences Program.

Ph.D. On-campus
5 years$32,776/yr
Columbia University4,4(113)
Manhattan, United States Top 0.5% in Worldwide

Neurobiology and Behavior

The Neurobiology and Behavior Program from Columbia University offer a diverse set of research and academic experiences that reflect the interdisciplinary nature of neuroscience. 

Ph.D. On-campus
7 years$56,840/yr

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