Overview

Modern biological research requires creative synergy between different disciplines. The TETRAD Graduate Program from University of California, San Francisco's curriculum provides students with a strong foundation in the basic concepts of biochemistry, genetics, developmental biology and cell biology through interactive courses. In the spring quarter, students choose three two-week long, discussion-based mini-courses that focus on a particular problem and encourage students to synthesize concepts from different disciplines to tackle the problem.

Career Outcomes

Students in the Tetrad Graduate Program have an excellent record of career success after graduation. The majority of graduates go on to either postdoctoral research positions or research-related or research-intensive careers

Programme Structure

Courses include:

Structure of Macromolecules

Cell Biology

Racism in Science

Biological Regulatory Mechanisms

Principles in Molecular Genetics

Minicourses

Key information

Start dates & application deadlines

Starting 2027-09-24 00:00:00
- Apply before 2026-12-01 00:00:00

Language

English

TOEFL^® IBT

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IELTS

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Delivered

On Campus

Campus Location

San Francisco, United States

Disciplines

Biology View 351 other PhDs in Biology in United States

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Academic requirements

GPA

3.2

English requirements

TOEFL^® IBT

Schedule TOEFL®

IELTS

Prepare for Your English Test

Cathoven AI IELTS Preparation

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AI-powered IELTS feedback. Clear, actionable, and tailored to boost your writing & speaking score. No credit card or upfront payment required.

Trusted by 300k learners
98 accuracy using real exam data
4.9/5 student rating

Other requirements

General requirements

Academics: We look carefully and holistically at the academic history and experience of each applicant, including any challenges you faced, which can be discussed in the personal statement.

Personal statement (AKA Statement of Purpose)

Comprehensive Research Statement: We consider prior research experience to be an important part of applying to our graduate program for two main reasons: 1) it helps us evaluate your potential as a researcher, and 2) it shows us that you have an understanding of how the intensive experience of graduate school aligns with your future career goals. Keep the research summary of each individual research experience brief (1-2 sentences) and use the Comprehensive Research Statement to tell us an overall story about your prior research experience, whether it be in academic labs, industry, or elsewhere.

Reference letters: We request you submit three confidential letters of reference on your behalf.

Tuition Fees

Tuition fees are shown in and the most likely applicable fee is shown based on your nationality.

International Applies to you

Applies to you

Non-residents

69993 USD / year

≈ 69993 USD / year
Out-of-State

69993 USD / year

≈ 69993 USD / year
Domestic

Applies to you

In-State

25839 USD / year

≈ 25839 USD / year

Living costs

San Francisco

United States

2206 - 4119 USD / month

The living costs include the total expenses per month, covering accommodation, public transportation, utilities (electricity, internet), books and groceries.

Funding

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Scholarships Information

Below you will find PhD's scholarship opportunities for TETRAD.

Available Scholarships

You are eligible to apply for these scholarships but a selection process will still be applied by the provider.

About

Overview

Career Outcomes

Programme Structure

Key information

Start dates & application deadlines

Language

Prepare for Your English Test

Delivered

Campus Location

Disciplines

What students do after studying

Academic requirements

English requirements

Prepare for Your English Test

Other requirements

General requirements

Tuition Fees

International Applies to you

Domestic

Living costs

San Francisco

Funding

Scholarships Information

Available Scholarships

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