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Integrative Biology, Ph.D.

  • Application Deadline
  • 48 months
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This unique multi-dimensional approach underpins our graduate program, where students combine observational, experimental, and comparative approaches with the development of theory, and apply concepts and techniques from the biological sciences and other disciplines.

Detailed Programme Facts

  • Programme intensity Full-time
    • Full-time duration 48 months
  • Languages
    • English
  • Delivery mode
    On Campus

Programme Structure

Courses include:

  • Principles of Phylogenetics: Systematics
  • Principles of Phylogenetics: Ecology and Evolution
  • Research Reviews in Animal Behavior: Behavior Review
  • Quantitative Methods for Ecological and Environmental Modeling
  • Integrative Biology Research Review
  • Seminar in Locomotion Energetics and Biomechanics
  • Seminar in Physiological Bases of Physical Activity
  • Seminar in Motor Control
  • Seminars in Isotopics
  • Stable Isotope Ecology
  • Marine Science Review
  • Seminar in Biomechanics
  • Seminar on Biology of Amphibians and Reptiles
  • Advanced Studies in Morphology
  • Endocrine Disruptors
  • Functional Neuroanatomy
  • IFunctional Neuroanatomy Laboratory
  • Seminar on Controversies in Comparative Physiology
  • Comparative Physiology and Endocrinology Seminar
  • Seminar on Evolutionary Genetics
  • Seminar in Ecology
  • Ecological Research Reviews

English Language Requirements

  • Minimum required score:


    The TOEFL iBT ® measures your English-language abilities in an academic setting. The test has four sections (reading, listening, speaking, and writing), each with a score range of 0-30, for a total score range of 0-120.

  • Minimum required score:


    The IELTS – or the International English Language Test System – tests your English-language abilities (writing, listening, speaking, and reading) on a scale of 1.00–9.00. The minimum IELTS score requirement refers to which Overall Band Score you received, which is your combined average score. Read more about IELTS.

    Get a free IELTS practice test
  • Minimum required score:


    The TOEFL®PBT is administered in a paper format and measures your ability to use and understand English in a classroom setting at the college or university level. It accurately measures how well you can listen, read and write in English while performing academic tasks.

General Requirements

Admissions Criteria
  • Students admitted to the program typically have a bachelor's degree in one of the life sciences or physical sciences. However, promising students with other academic backgrounds are encouraged to apply if they have a undergraduate grounding in biology.
  • Grade Point Average (GPA) - Upper division or graduate GPA of 3.4 or higher is preferred. A minimum GPA of 3.0 (courses taken after the first two years) is required by the Graduate Division.
  • Three letters of recommendation from faculty or other persons who have known you in an academic or research capacity. 
  • Statement of Purpose

  • Personal History Statement

Tuition Fee

  • International

    33958 USD/year
    Tuition Fee
    Based on the original amount of 16979 USD per semester and a duration of 48 months.
  • National

    33958 USD/year
    Tuition Fee
    Based on the original amount of 16979 USD per semester and a duration of 48 months.
  • In-State

    18856 USD/year
    Tuition Fee
    Based on the original amount of 9428 USD per semester and a duration of 48 months.
We've labeled the tuition fee that applies to you because we think you are from and prefer over other currencies.

Living costs for Berkeley

  • 1750 - 2820 USD/month
    Living Costs

The living costs include the total expenses per month, covering accommodation, public transportation, utilities (electricity, internet), books and groceries.


Studyportals Tip: Students can search online for independent or external scholarships that can help fund their studies. Check the scholarships to see whether you are eligible to apply. Many scholarships are either merit-based or needs-based.


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